Rosemary G. Gillespie is a professor and the Schlinger Chair in Systematic Entomology in UC Berkeley's Department of Environmental Science, Policy, and Management. She is president elect of the American Genetics Association, past president of the International Biogeography Society, trustee and fellow of the California Academy of Sciences, senior editor for Molecular Ecology, and associate editor for Journal of Biogeography. Her primary research uses islands as model systems to understand ecological and evolutionary processes. Hotspot archipelagoes, such as Hawaii, provide a temporal framework of islands that allows one to synthesize ecological and evolutionary perspectives. She uses this framework to integrate macroecological (interaction networks and maximum entropy inference) and evolutionary (population genetics and phylogenetics) approaches to build a predictive understanding of the dynamic interplay between ecology and evolution in shaping the ecology of complex ecosystems.